duke university biology phd programs

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Graduate program.

Duke Biology offers diverse training opportunities for students seeking a Ph.D. degree in biological sciences. Our program is highly flexible, allowing students to design the path that best fits their professional needs. Our students spend the majority of their time engaged in innovative research, and have the opportunity to interact and collaborate with researchers from a wide range of fields. 

The graduate students in the Biology Department (Biograds) are integral to the department and community. Contributions include: running weekly happy hours and prospective student recruitment weekends as well as organizing seminars, journal clubs, teaching-assistant training orientation, and diversity-themed workshops.

Program Requirements

Requirements for the Ph.D. include course work, participation in seminars relevant to your field of research, successful completion of an oral qualifying examination, one year of teaching experience, completion and defense of a research dissertation under faculty guidance.

Affiliated Graduate Programs

Duke Biology is involved in multiple University-wide graduate programs, which link researchers across different departments and schools at Duke – most notably the basic sciences departments in the  Duke University Medical Center  and the  Nicholas School of Earth and Ocean Sciences .

For Prospective Students

Learn more about our degree, our university, and our community. We'll explain the application process, financial support, and more.

• For Current Students

We've compiled resources to assist you while working toward your Ph.D. – from requirements and fellowships, to milestones, useful Duke links, and more.

Learn more about the application process, deadlines and other resources for our candidates.

Review Application Information

• Duke Biology’s Mission Statement
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• Learning from Baboons: Dr. Susan Alberts
• Extremophiles and Systems Biology: Dr. Amy Schmid
• How Cells Manage Stress: Dr. Gustavo Silva
• Predator-Prey Interactions in a Changing World: Dr. Jean Philippe Gibert
• Exploring the Extracellular Matrix: Dr. David Sherwood
• Cell Division's Missing Link: Dr. Masayuki Onishi
• Listening in to Birdsong: Dr. Steve Nowicki
• Biogeochemistry as Ecosystem Accounting: Dr. Emily Bernhardt
• Building a Dynamic Nervous System: Dr. Pelin Volkan
• Investigating a Key Plant Hormone: Dr. Lucia Strader
• Imagining Visual Ecology: Dr. Sönke Johnsen
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G-BIO-PHD - Biology - PhD

Degree designation.

The Department of Biology offers a variety of training opportunities leading to the PhD in biology.

Students in the department may specialize in a wide variety of areas including anatomy; behavior; physiology; cellular and molecular biology; community, ecosystem, physiological, and population ecology; evolution; functional morphology; developmental, ecological, molecular, organelle, and population genetics; genomics; and phylogenetic systematics.

There is a high level of interaction among the various areas of biology and other programs. Faculty members participate in the University Programs in Developmental Biology, Ecology, Genetics and Genomics, Cellular and Molecular Biology, Computational Biology and Bioinformatics, Structural Biology and Biophysics, and Neurobiology; tropical research is facilitated through the university’s membership in the Organization for Tropical Studies. There are also strong relationships with the Departments of Evolutionary Anthropology (primatology, phylogenetic systematics, macroevolution), Mathematics (theoretical biology), and Psychology (behavior); the Pratt School of Engineering (biomechanics); the Medical Center (molecular biology and genomics); and the Nicholas School of the Environment (ecology).

Students entering the program generally have a broad background in biological sciences supplemented with basic courses in chemistry, mathematics, and physics. Biochemistry and physical chemistry are strongly recommended for students interested in molecular areas, and advanced courses in mathematics are recommended for students in population genetics and ecology. While deficiencies may be corrected by taking appropriate courses during the first year of graduate study, it is advised that students search widely in this bulletin for information about the intellectual resources of the university. Courses below the 500 level may not be applied toward the required credits needed for a post-baccalaureate degree. With the approval of their Director of Graduate Studies and the Associate Dean for Academic Affairs, graduate students may enroll in lower-level courses, but these courses will not count toward any graduation requirement and will not be included in a student’s GPA calculation. Special attention should be given to announcements of the programs and departments listed above, as well as to those of Cultural Anthropology, History, Immunology, Molecular Genetics and Microbiology, Pharmacology, Philosophy, and Sociology, and of the Pratt School of Engineering and the Nicholas School of the Environment.

Ph.D. Programs

Biological and biomedical sciences, physical sciences and engineering, social sciences.

* – Denotes Ph.D. admitting programs. Students may apply and be admitted directly to these departments or programs, but the Ph.D. is offered only through one of the participating departments identified in the program description. After their second year of study at Duke, students must select a participating department in which they plan to earn the Ph.D.

Biochemistry Biology Biostatistics Cell and Molecular Biology Cell Biology Cognitive Neuroscience* Computational Biology and Bioinformatics Developmental and Stem Cell Biology* Ecology Evolutionary Anthropology Genetics and Genomics

Immunology Integrated Toxicology and Environmental Health* Medical Physics Medical Scientist Training Molecular Cancer Biology Molecular Genetics and Microbiology Neurobiology Pathology Pharmacology Population Health Sciences

Art, Art History and Visual Studies Classical Studies Computational Media, Arts & Cultures English German Studies (Carolina-Duke German Program)

Literature Music Philosophy Religious Studies Romance Studies

Biomedical Engineering Chemistry Civil and Environmental Engineering Computer Science Earth and Climate Sciences Electrical and Computer Engineering Environment

Marine Science and Conservation Materials Science and Engineering Mathematics Mechanical Engineering and Materials Science Physics Statistical Science

Business Administration Cultural Anthropology Economics Environmental Policy History

Nursing Political Science Psychology and Neuroscience Public Policy Sociology

Applying to the Graduate Program in Cell Biology

The Department of Cell Biology offers a program of study leading to the PhD. degree in Cell Biology. The goal of this program is to train students in cell and developmental biology to become independent, innovative and critical scientists fully prepared for productive careers in academic and other settings. The Department offers strengths in neuronal, molecular, developmental and stem cell biology.

Admission to the Program

The Cell Biology Department does not accept applications directly. Rather, students enter Cell Biology from any of the interdepartmental training programs:

Cell and Molecular Biology ,

the University Program in Genetics

the Structural Biology and Biophysics Program

Molecular Cancer Biology

Department of Molecular Genetics and Microbiology

and the Developmental and Stem Cell Biology Program .

Students interested in the research programs in Cell Biology should apply to the Graduate School , listing one of these programs. Admission to one of these programs guarantees funding for the term of the Ph.D. Some funding slots are available to foreign students. In the first year, a student should plan to do a lab rotation with any Cell Biology faculty of interest. All students who decide to do thesis work with a Cell Biology faculty member are accepted into the department, and maintain dual association with their interdepartmental program.

All students seeking to affiliate with a Cell Biology lab must send to DGS, along with the required forms, a brief (200 words max.) description of project and justification of the choice of lab. Students affiliating with a lab whose PI's primary affiliation is not Cell Biology must also add a brief (100 word max) justification for choosing the Cell Biology Graduate Program.  

Lab Rotations and Choice of a Thesis Lab

Each first-year student is expected to complete a series of 3 laboratory rotations.

The student may choose the labs, subject to approval by laboratory directors;

Each rotation involves two to three months of work: the first rotation usually runs from September through November, the second from December through February, and the third from March through May.

In general, students are expected to spend about 15 hours weekly in laboratory work. We encourage students to discuss this expectation with the principal investigator before beginning the rotation.

At the end of each rotation, the student should expect to give a presentation to the laboratory director and/or prepare a brief report summarizing experiments and results. By offering detailed exposure to procedures and techniques in three separate lab settings, these rotations provide the basis for choosing a thesis laboratory.

Following the third rotation, a student selects a laboratory for thesis work and begins a research program there. The choice of thesis laboratory must be approved by the thesis advisor (who must have the funding, lab space, & resources to support the research) and by the Director of Graduate Studies.  Forms needed to affiliate with your new laboratory can be obtained from your admitting program DGSA .

 All students seeking to affiliate with a Cell Biology lab must send to DGS along with the required forms a brief (200 words max.) description of project and justification of the choice of lab. Students affiliating with a lab whose PI's primary affiliation is not Cell Biology must also add a brief (100 word max) justification for choosing the Cell Biology Graduate Program.  

Recruitment Opportunities:

The following faculty members in Cell Biology are currently recruiting Graduate Students.  Please click on the faculty's name to read more about their research: Michel Bagnat Blanche Capel Yarui Diao Stefano Di Talia Cagla Eroglu Chantell Evans Akankshi Munjal Chris Nicchitta Ken Poss Scott Soderling Purushothama Rao Tata Eda Yildirim

Have feedback about our website? Can't find what you're looking for? Need to report a broken link? Click here.

Our PhD training program in Developmental and Stem Cell Biology (DSCB) is the culmination of a long tradition of excellence in training developmental biologists at Duke University. Recent technological and conceptual advances have ushered in an era of unprecedented opportunities to uncover key mechanisms of embryogenesis, stem cell biology, regeneration and other developmental phenomena. The DSCB Program is central to this research enterprise at Duke, and provides a superb training environment.

The mission of the DSCB PhD program  is to train the next generation of scientists in principles of development, stem cell biology, and regeneration through targeted coursework, lab research, and interactions within the DSCB community. The diverse backgrounds, research expertise, and perspectives of our community enable us to enrich and personalize the training environment. Our goal is to develop in our students confidence, independent critical thinking, academic rigor, and the intellectual curiosity and creativity to synthesize new ideas and communicate them effectively.

Admissions        Diversity        DSCB Faculty    

Recent News

The Pharmacology program at Duke is consistently ranked among the top pharmacology graduate programs in the nation. Its focus is to prepare qualified individuals for a career in independent research. Pharmacology is the science of drug action on biological systems. It encompasses the study of targets of drug action, the mechanisms by which drugs act, the therapeutic and toxic effects of drugs, as well as the development of new therapeutic agents. As the study of pharmacology is interdisciplinary, the graduate program in pharmacology is diverse and flexible. Graduate positions in the program are fully funded providing for payment of tuition, fees, and an annual stipend for the first two years. After the first two years, students are supported by the faculty member with whom they are doing their thesis research. The average time to completion of a Ph.D. is 5.5 years.

Pharmacology Alumni

We are very proud of the alumni from our department who have successfully completed their training in Pharmacology. Please follow this link to learn more about our alumni for   pharmacology .

David MacAlpine, PhD

McDonnell inducted into the Royal Irish Academy

• Oliver Lab finds that olfactory neuroblastoma smells a lot like small cell lung cancer in Cancer Cell

• Wang Lab identifies a new player in regulating neuronal-immune interactions in Nature Immunology

• Dr. Lee Zou is the recipient of the George Barth Geller Distinguished Professorship

Recent News

Olfactory neuroblastoma smells a lot like small cell lung cancer

A new player in regulating neuronal-immune interactions in the context of cancer immunotherapy identified by Wang Lab

Education & Training

Upcoming Events

Graduate School

Biochemistry (ph.d.), biochemistry (ph.d.) | graduate.

Our Biochemistry doctoral students are at the forefront of biochemical research and molecular medicine, examining biological mechanisms underlying human disease; they are finding new ways to detect and attack diseases and immunological disorders like cancer, neurological disorders, and cardiovascular disease.

Graduates of the Ph.D. in Biochemistry program at Howard's Graduate School are prepared for careers at top research universities and senior-level research positions in biomedical and related industries. The program's key strengths in molecular microbiology, proteomics and genetics, bioinformatics, and drug design and discovery make us a nexus for collaborative investigations between biochemistry researchers and clinicians. You'll learn to apply biochemical techniques, including NMR spectroscopy, crystallography, and single-molecule methods as well as contemporary approaches to cell culture and genetic analysis to answer key questions about the pathogenesis of specific diseases and the development of effective drug therapies. You'll also enjoy the close mentorship of faculty who are committed to your professional development. Our faculty are experts in several areas of biochemistry, including analysis of molecular structure, proteomics and genetics, tumor biology, structural biology, enzymology, RNA catalysis, stress response, and RNA modification. As you advance in the program, you'll become increasingly involved in laboratory research and the critical analysis of biochemical literature. Our graduate seminar series offers a venue to present your early-stage research. Students may pursue a dual M.D./Ph.D. degree.

Program Snapshot

      ❱  72 credit hours        ❱  Full-time       ❱  On-campus format       ❱  Degree: Ph.D.       ❱  Dual degree: M.D./Ph.D.

Application Deadlines

Spring 2024 entry:         ❱  No spring entry 

Fall 2024 entry:         ❱  Dec. 1, 2023 (early deadline)       ❱  Feb. 15, 2024 (priority deadline)       ❱  Apr. 15, 2024 (final deadline)

Applicants should submit their applications as early as possible for earlier consideration of departmental funding opportunities. Applicants have until the final deadline to apply. However, applications will be reviewed on a rolling basis throughout the admissions cycle. 

Dr. Zaki Sherif

Dr. matthew george, jr., angela wilson, program details.

• Degree Classification: Graduate
• Related Degrees: M.D. / Ph.D., Ph.D.

Admission Requirements

Application for admission.

• Online GradCAS application
• Statement of purpose/ Statement of academic interest ( 500-1,000 words )
• GRE scores  not required
• Official transcripts sent to GradCAS
• 3 letters of recommendation
• Bachelor's degree from an accredited college or university or the international equivalent 
• Resume or Curriculum Vitae
• Autobiographical statement ( 500-750 words )

GRE Required?

Gre preferred minimums.

• GRE Verbal Reasoning: N/A
• GRE Quantitative Reasoning: N/A
• GRE Analytical Writing: N/A

GPA Required Minimums

• Overall GPA minimum: 3.0
• Undergrad GPA minimum: 3.0

Prerequisite Courses

The following course prerequisites are required. Applicants are required to have at least a B average in these prerequisites.   No expiration date for recommended prerequisites.

• Biology (college-level courses, 8 semester credit hrs)
• General Chemistry (college-level courses, 8 semester credit hrs)
• Organic Chemistry (college-level courses, 8 semester credit hrs)
• Elementary Physical Chemistry (college-level course and lab, 4 semester credit hrs)
• Physics (college-level courses, 8 semester credit hrs)
• Calculus  (college-level course, 3 semester credit hrs)

Reference Requirements

Evaluator type accepted:

• Professor (Required)
• Supervisor/Manager
• Other 

Evaluator type not accepted:

• Family Member

40 Facts About Elektrostal

Written by Lanette Mayes

Modified & Updated: 21 May 2024

Reviewed by Jessica Corbett

Elektrostal is a vibrant city located in the Moscow Oblast region of Russia. With a rich history, stunning architecture, and a thriving community, Elektrostal is a city that has much to offer. Whether you are a history buff, nature enthusiast, or simply curious about different cultures, Elektrostal is sure to captivate you.

This article will provide you with 40 fascinating facts about Elektrostal, giving you a better understanding of why this city is worth exploring. From its origins as an industrial hub to its modern-day charm, we will delve into the various aspects that make Elektrostal a unique and must-visit destination.

So, join us as we uncover the hidden treasures of Elektrostal and discover what makes this city a true gem in the heart of Russia.

Key Takeaways:

• Elektrostal, known as the “Motor City of Russia,” is a vibrant and growing city with a rich industrial history, offering diverse cultural experiences and a strong commitment to environmental sustainability.
• With its convenient location near Moscow, Elektrostal provides a picturesque landscape, vibrant nightlife, and a range of recreational activities, making it an ideal destination for residents and visitors alike.

Known as the “Motor City of Russia.”

Elektrostal, a city located in the Moscow Oblast region of Russia, earned the nickname “Motor City” due to its significant involvement in the automotive industry.

Home to the Elektrostal Metallurgical Plant.

Elektrostal is renowned for its metallurgical plant, which has been producing high-quality steel and alloys since its establishment in 1916.

Boasts a rich industrial heritage.

Elektrostal has a long history of industrial development, contributing to the growth and progress of the region.

Founded in 1916.

The city of Elektrostal was founded in 1916 as a result of the construction of the Elektrostal Metallurgical Plant.

Located approximately 50 kilometers east of Moscow.

Elektrostal is situated in close proximity to the Russian capital, making it easily accessible for both residents and visitors.

Known for its vibrant cultural scene.

Elektrostal is home to several cultural institutions, including museums, theaters, and art galleries that showcase the city’s rich artistic heritage.

A popular destination for nature lovers.

Surrounded by picturesque landscapes and forests, Elektrostal offers ample opportunities for outdoor activities such as hiking, camping, and birdwatching.

Hosts the annual Elektrostal City Day celebrations.

Every year, Elektrostal organizes festive events and activities to celebrate its founding, bringing together residents and visitors in a spirit of unity and joy.

Has a population of approximately 160,000 people.

Elektrostal is home to a diverse and vibrant community of around 160,000 residents, contributing to its dynamic atmosphere.

Boasts excellent education facilities.

The city is known for its well-established educational institutions, providing quality education to students of all ages.

A center for scientific research and innovation.

Elektrostal serves as an important hub for scientific research, particularly in the fields of metallurgy , materials science, and engineering.

Surrounded by picturesque lakes.

The city is blessed with numerous beautiful lakes , offering scenic views and recreational opportunities for locals and visitors alike.

Well-connected transportation system.

Elektrostal benefits from an efficient transportation network, including highways, railways, and public transportation options, ensuring convenient travel within and beyond the city.

Famous for its traditional Russian cuisine.

Food enthusiasts can indulge in authentic Russian dishes at numerous restaurants and cafes scattered throughout Elektrostal.

Home to notable architectural landmarks.

Elektrostal boasts impressive architecture, including the Church of the Transfiguration of the Lord and the Elektrostal Palace of Culture.

Offers a wide range of recreational facilities.

Residents and visitors can enjoy various recreational activities, such as sports complexes, swimming pools, and fitness centers, enhancing the overall quality of life.

Provides a high standard of healthcare.

Elektrostal is equipped with modern medical facilities, ensuring residents have access to quality healthcare services.

Home to the Elektrostal History Museum.

The Elektrostal History Museum showcases the city’s fascinating past through exhibitions and displays.

A hub for sports enthusiasts.

Elektrostal is passionate about sports, with numerous stadiums, arenas, and sports clubs offering opportunities for athletes and spectators.

Celebrates diverse cultural festivals.

Throughout the year, Elektrostal hosts a variety of cultural festivals, celebrating different ethnicities, traditions, and art forms.

Electric power played a significant role in its early development.

Elektrostal owes its name and initial growth to the establishment of electric power stations and the utilization of electricity in the industrial sector.

Boasts a thriving economy.

The city’s strong industrial base, coupled with its strategic location near Moscow, has contributed to Elektrostal’s prosperous economic status.

Houses the Elektrostal Drama Theater.

The Elektrostal Drama Theater is a cultural centerpiece, attracting theater enthusiasts from far and wide.

Popular destination for winter sports.

Elektrostal’s proximity to ski resorts and winter sport facilities makes it a favorite destination for skiing, snowboarding, and other winter activities.

Promotes environmental sustainability.

Elektrostal prioritizes environmental protection and sustainability, implementing initiatives to reduce pollution and preserve natural resources.

Home to renowned educational institutions.

Elektrostal is known for its prestigious schools and universities, offering a wide range of academic programs to students.

Committed to cultural preservation.

The city values its cultural heritage and takes active steps to preserve and promote traditional customs, crafts, and arts.

Hosts an annual International Film Festival.

The Elektrostal International Film Festival attracts filmmakers and cinema enthusiasts from around the world, showcasing a diverse range of films.

Encourages entrepreneurship and innovation.

Elektrostal supports aspiring entrepreneurs and fosters a culture of innovation, providing opportunities for startups and business development.

Offers a range of housing options.

Elektrostal provides diverse housing options, including apartments, houses, and residential complexes, catering to different lifestyles and budgets.

Home to notable sports teams.

Elektrostal is proud of its sports legacy, with several successful sports teams competing at regional and national levels.

Boasts a vibrant nightlife scene.

Residents and visitors can enjoy a lively nightlife in Elektrostal, with numerous bars, clubs, and entertainment venues.

Promotes cultural exchange and international relations.

Elektrostal actively engages in international partnerships, cultural exchanges, and diplomatic collaborations to foster global connections.

Surrounded by beautiful nature reserves.

Nearby nature reserves, such as the Barybino Forest and Luchinskoye Lake, offer opportunities for nature enthusiasts to explore and appreciate the region’s biodiversity.

Commemorates historical events.

The city pays tribute to significant historical events through memorials, monuments, and exhibitions, ensuring the preservation of collective memory.

Promotes sports and youth development.

Elektrostal invests in sports infrastructure and programs to encourage youth participation, health, and physical fitness.

Hosts annual cultural and artistic festivals.

Throughout the year, Elektrostal celebrates its cultural diversity through festivals dedicated to music, dance, art, and theater.

Provides a picturesque landscape for photography enthusiasts.

The city’s scenic beauty, architectural landmarks, and natural surroundings make it a paradise for photographers.

Connects to Moscow via a direct train line.

The convenient train connection between Elektrostal and Moscow makes commuting between the two cities effortless.

A city with a bright future.

Elektrostal continues to grow and develop, aiming to become a model city in terms of infrastructure, sustainability, and quality of life for its residents.

In conclusion, Elektrostal is a fascinating city with a rich history and a vibrant present. From its origins as a center of steel production to its modern-day status as a hub for education and industry, Elektrostal has plenty to offer both residents and visitors. With its beautiful parks, cultural attractions, and proximity to Moscow, there is no shortage of things to see and do in this dynamic city. Whether you’re interested in exploring its historical landmarks, enjoying outdoor activities, or immersing yourself in the local culture, Elektrostal has something for everyone. So, next time you find yourself in the Moscow region, don’t miss the opportunity to discover the hidden gems of Elektrostal.

Q: What is the population of Elektrostal?

A: As of the latest data, the population of Elektrostal is approximately XXXX.

Q: How far is Elektrostal from Moscow?

A: Elektrostal is located approximately XX kilometers away from Moscow.

Q: Are there any famous landmarks in Elektrostal?

A: Yes, Elektrostal is home to several notable landmarks, including XXXX and XXXX.

Q: What industries are prominent in Elektrostal?

A: Elektrostal is known for its steel production industry and is also a center for engineering and manufacturing.

Q: Are there any universities or educational institutions in Elektrostal?

A: Yes, Elektrostal is home to XXXX University and several other educational institutions.

Q: What are some popular outdoor activities in Elektrostal?

A: Elektrostal offers several outdoor activities, such as hiking, cycling, and picnicking in its beautiful parks.

Q: Is Elektrostal well-connected in terms of transportation?

A: Yes, Elektrostal has good transportation links, including trains and buses, making it easily accessible from nearby cities.

Q: Are there any annual events or festivals in Elektrostal?

A: Yes, Elektrostal hosts various events and festivals throughout the year, including XXXX and XXXX.

Elektrostal's fascinating history, vibrant culture, and promising future make it a city worth exploring. For more captivating facts about cities around the world, discover the unique characteristics that define each city . Uncover the hidden gems of Moscow Oblast through our in-depth look at Kolomna. Lastly, dive into the rich industrial heritage of Teesside, a thriving industrial center with its own story to tell.

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